This, like several other species that occur in isolated regions of southern France, is principally an Iberian species which "leaks" into the extreme south of France, where it is highly localised. It also occurs in southern Italy and southern Greece. The upperside is superficially similar to Pyrgus grizzled skippers although it is not of the Pyrgus genus. The underside is quite different, the markings being more diffuse than the clearly delineated marks of Pyrgus species, and having more in common with Carcharodus species. It is quite large, perhaps rather larger than the largest of the Pyrgus species.

It is generally a species of medium altitudes, with a range of 0-1000m. It is univoltine from June to October, although most common in late summer.

It is generally found in the vicinity of its larval hostplant, various species of the Phlomis genus, the purple-flowered P. herba-venti (as shown in 54105), the yellow-flowered P. lychnitis, or (according to Lafranchis), in the PACA region, P. fruticosa . In the two locations where proto has been seen in Var, P. herba-venti was common. However, in the one location I have seen proto in Var in the greatest numbers, it appeared to be in a region of P. fruticosa.

In 2025 I visited a site in the Cévennes and found a small colony of proto in a meadow where P. herba-venti was growing.

The late August and September photographs on this page are from the end of the flight period where most of the twenty or so individuals seen in 2014 were showing signs of wear, with only a few females still looking reasonably fresh. They were nectaring incessantly, so these photographs were snapped on the move. I revisited the location in mid-July 2017 which was clearly the start of the flight period, as only males were flying and they were mostly quite fresh. They were nectaring on white Scabious (is this Field Scabious (Knautia arvensis)) - can this be white as well as lilac or pink? In 2019, a notable late season, I visited the same site in Var in mid-July and proto had not yet emerged.

In 2022 and 2023 excessive temperatures and absence of rainfall for long periods created a very baked and flower-free terrain in southern Provence; very few butterflies were on the wing in late August, but proto was still going reasonably strong.

The English name is something of a misnomer in my experience, as most are decidedly brown and only 45673 could be described as sage green from whence it presumably gets its name.

This species was previously known as Syrichtus proto. 

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